← Dynamical Systems in Biology

Jacobian matrices

The Jacobian matrix collects the first partial derivatives of a multidimensional dynamical system.

Two-variable system

If

\[\frac{dx}{dt}=f(x,y),\qquad \frac{dy}{dt}=g(x,y),\]

then

\[J(x,y)=\begin{pmatrix}\frac{\partial f}{\partial x}&\frac{\partial f}{\partial y}\\[4pt]\frac{\partial g}{\partial x}&\frac{\partial g}{\partial y}\end{pmatrix}.\]

Evaluation at an equilibrium

At an equilibrium \((x^*,y^*)\), the matrix \(J(x^*,y^*)\) describes how small changes in the state alter the local rates of change.

Biological interpretation

Each entry measures the local sensitivity of one equation to one state variable. Off-diagonal terms often reflect interactions between biological components.

Key idea. The Jacobian is the local derivative of the full vector field and is the central matrix used in linear stability analysis.